B.S., 1981, The University of Michigan, Ann Arbor; M.S. 1984, The University of Michigan, Ann Arbor; Ph.D., 1988, The University of Michigan, Ann Arbor; 1988 Postdoctoral Fellow, Chalmers University of Technology, Sweden; 1989-91 Postdoctoral Fellow, The University of Michigan, Ann Arbor; 1991-94 Senior Research Fellow, Mayo Clinic.